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The main of these changes can be characterized as follows:
Alexander also lost muscle fibers completely, becoming a ligament (morphological and functional substitution), the following muscles, functioning in remote polydactyl ancestors of modern horses:
1. In the elbow joint - m. pronator teres, turned into a long branch of the medial collateral ligament of the joint, and possibly m. supinator longus 2), if that was considered a strong oblique dorsal strand, which is closely connected with the fibrous layer of the dorsal surface of the capsule of elbow joint (ligam. obliquum); final division of their fibers is partially fused with m. pronator teres h).
2. In the radioulnar joint at the level of the proximal interosseous gap is converted into a medial transverse ligament of the radius and ulna bones m. pronator quadratus, reinforcing the mutual attachment of proximal epiphyses of both forearm bones horse.Thus, the loss of the radius and ulna bones of the mutual mobility of the muscles, providing in the past, this mobility, survived, but their elements replaced the muscle connective tissue and ligaments have become used animals to strengthen the elbow and strengthening its spring properties (articular connection between the proximal endsboth forearm bones in a horse, usually stored).
3. In the role of powerful brushes the proximal ligament sesamoid bones of the first phalanx (ligam. suspensoriuin ossium sesamoideorum phalangis primae) acquires a horse's third interosseous muscle - m. interosseus medius). Losing value interosseous muscle, the muscle to maintain and strengthen its ties with the finger dorsal aponeurosis and the latter acquired through close contact with the final Department of finger extensor tendons of the total (m. extensor digitalis communis), formed for this tendon two auxiliary heads of the intermediate mounting sssamovidnyh bones of the 1 st phalanx. In addition, m. interosseus medius extended its proximal attachment on the surface of the distal volyarnuyu number of bones and ligaments deep carpal joint.
Thus, the finger joints are placed in close functional relationship of the carpal joint.
4. In thin tendon heavy, combining both activities of finger extensor forelimb and ensure close liaison between the tendons have become their own second finger extensor tendon and the total finger extensor muscle of the fourth finger in the form of so-called muscle Tirnessa and Phillips.
5. In the tibia (dorsal) placed turned into a tendinous band sh. peronaeus (s. frbularis) tertius, which is also called "tendo femoro-tarseus". In the past, this is definitely a muscle, having lost in the process of specialization of their muscle fibers and to establish a close relationship with the connective tissue surrounding muscle divisions.
-6. The same can be said about the second "muscle", located on the opposite (plantar) surface of the tibia, the so-called co. Flexor digitalis pedis sublimis (s. superficialis), which is still called "tendo plantaris".
This specialized arm has so little muscle fibers that affect the change of its length, they can not, they only reinforce its elasticity.
7. Is also important to establish a correlative relationship between the knee and joints tarsalnym number of flat strands of connective tissue located plantar to calf.